Coleus scutellarioides

(L.) Benth. (1830)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Lamiales Family: Lamiaceae Genus: Coleus

coleus, coleus, weleweka, weleweka

Description

Key Characters:

Growth Form: Erect perennial herbs.

Stems: Stems fleshy, 5–15 dm long, sparsely to moderately puberulent.

Roots:

Leaves: Leaves simple. Opposite. Blades ovate–deltate to broadly ovate, (7–)10–21 cm long, 2.5–10 cm wide. Upper surfaces sparsely to densely scabrid to hirtellous or strigillose; lower surfaces glandular punctate and hirtellous, especially on veins, Leaves variously colored green, red, purple, yellow, or variegated, membranous. Margins crenate to serrate. Petioles 2–5.5 cm long, strigillose. Stipules absent.

Flowers: Flowers in verticillasters or irregularly branched cymes 5–10(–25) cm long, bracts ovate, caducous; bracts early deciduous or absent; flowers pedicellate. Calyx actinomorphic, occasionally bilabiate, 2–2.5 mm long, enlarging to 4–6 mm long in fruit, hirtellous and glandular punctate, the glands black, upper lip broadly ovate, becoming recurved in fruit, apex acute, lower lip 4-toothed, the teeth distinctly unequal, lateral ones very short, truncate to rounded, median ones sharply pointed. Corolla zygomorphic, much exserted, blue or violet, tube whitish, lower lip boat–shaped, longer than upper lip, 8–13 mm long; nectary disk often present at base of ovary, annular or developed on anterior side only. Stamens 4, declinate, nearly included in corolla tube; filaments connate or free; anther cells apex confluent, dithecal, opening by longitudinal slits. Ovary superior, 2-carpellate, placentation basal-axile; ovules 1 per ovary lobe, anatropous to hemitropous or apotropous; style 1, apex equally 2-cleft, shortly 2 lobed; stigmas 2.

Fruit: Nutlets 1-seeded; brown; shiny; broadly ovoid to lenticular; 1–1.2 mm long. Seeds with endosperm absent or scanty and oily.

Ploidy: 2n = 16; 24; 36; 40; 48; 49; 54; 72

Habitat: Various forms are cultivated; some of which are now naturalized in disturbed parts of mesic to wet forest.

Elevation Range: 30–300 m.

Historical Distribution

Synonyms (101)

Uses and Culture

USES

  • Yes

Natural History

Statewide Status

Naturalized

Island Status

Kaua'i Naturalized
O'ahu Naturalized
Maui Naturalized
Hawai'i Naturalized

Dispersal Agents


Pollinators

Specimens

Bibliography

Name Published In: N.Wallich, Pl. Asiat. Rar. 2: 16 (1830)

Other References

Wagner et al. 1990:828 (K, O, M, H [as Plectranthus scutellarioides]); Wagner & Herbst 1999:1883 (NOM:Syn. P. scutellarioides = Solenostemon scutellarioides); Staples & Herbst 2005:356 (KEY), 364 (DESCR); Grayer et al. 2010/Wagner et al. 2012:48 (NOM:Syn. S. scutellarioides)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Coleus scutellarioides (L.) Benth. Laie semi wet area PRESERVED_SPECIMEN Guinto, J. 22 Oahu BISH 3/6/1986
2 Coleus scutellarioides (L.) Benth. Tantalus, 3245 Round Top Dr, at Puu Ualakaa gate Mesic wet forest, disturbed roadside PRESERVED_SPECIMEN G. J. Ray 42 NY 8/27/1997
3 Coleus scutellarioides (L.) Benth. tantalus, 3245 round top dr, at puu ualakaa gate Mesic wet forest, disturbed roadside PRESERVED_SPECIMEN g. j. ray 42 NY 8/27/1997
4 Coleus scutellarioides (L.) Benth. hanalei valley, ohiki rd. PRESERVED_SPECIMEN t. w. flynn 1608 PH 3/24/1986
5 Coleus scutellarioides (L.) Benth. Limahuli Valley, National Tropical Botanical Garden. Cultivated plants. Cultivated. PRESERVED_SPECIMEN Tim Flynn 3765 Kauai PTBG 2/15/1990
6 Coleus scutellarioides (L.) Benth. kauai island: hanalei, wainiha power house road. next to power house Cultivated PRESERVED_SPECIMEN h. t. beck 1200 NY 8/8/1990
7 Coleus scutellarioides (L.) Benth. Hanalei Valley. Naturalized in pasture along Ohiki Road. PRESERVED_SPECIMEN Tim Flynn 1608 Kauai PTBG 3/24/1986
8 Coleus scutellarioides (L.) Benth. NaPali Coast State Park, back of Kalalau Valley. Next to swimming hole along Kalalau Stream. Degraded Diverse Mesic Forest. PRESERVED_SPECIMEN Natalia Tangalin 2382 Kauai PTBG 8/21/2010
9 Coleus scutellarioides (L.) Benth. KAUAI: Hanalei Valley. Naturalized in pasture along Ohiki Road. PRESERVED_SPECIMEN Tim Flynn 1608 Kauai PTBG 3/24/1986
10 Coleus scutellarioides (L.) Benth. Honomaele. Ula'ino road. PRESERVED_SPECIMEN Hank Oppenheimer H60120 Maui PTBG 6/11/2001
11 Coleus scutellarioides (L.) Benth. Kalahu Point. PTBG Kahanu Garden. Elev. 0-50 ft. Cultivated area around Piilanihale Heiau. In remnant Pandanus forest. PRESERVED_SPECIMEN Tim Flynn 2473 Maui PTBG 10/20/1987
12 Coleus scutellarioides (L.) Benth. stainback hwy. near jct. with hwy. 11. PRESERVED_SPECIMEN d. r. herbst 926 Hawaii US 2/16/1968
13 Coleus scutellarioides (L.) Benth. Hana, along Olopawa road to cinder pit. Alien veg. with Aleurities moluccana, Spathodea campanulata. PRESERVED_SPECIMEN Steve Perlman 22928 Maui PTBG 6/21/2012
14 Coleus scutellarioides (L.) Benth. Haiku Stairs, base of stairs near road. PRESERVED_SPECIMEN W. L. Wagner 5643 Oahu PTBG 6/2/1985
15 Coleus scutellarioides (L.) Benth. Kapaa, Kanepoonui Road off of Kawaihau Road. Collected as part of KISC 2007 Early Detection Roadside Survey. PRESERVED_SPECIMEN Natalia Tangalin 1318 Kauai PTBG 5/29/2007
16 Coleus scutellarioides (L.) Benth. Kalalau, just above Big Pools, planted next to the stream with Impatiens walleriana. PRESERVED_SPECIMEN Natalia Tangalin 2382 Kauai PTBG 8/23/2010
17 Coleus scutellarioides (L.) Benth. Honolulu: Manoa Valley; University of Hawaii Manoa Campus, courtyard of St.John Hall PRESERVED_SPECIMEN Lau, J. 1175 Oahu US 7/1/1985
18 Coleus scutellarioides (L.) Benth. Honolulu PRESERVED_SPECIMEN Brigham, W. T. s.n. Oahu US 3/4/1905
19 Coleus scutellarioides (L.) Benth. hamakua district, hana, along olopawa road to cinder pit. PRESERVED_SPECIMEN s. p. perlman, h. l. oppenheimer & k. m. bustamente 22928 Maui US 6/21/2012
20 Coleus scutellarioides (L.) Benth. Hamakua District, Hana, along Olopawa road to cinder pit. PRESERVED_SPECIMEN S. P. Perlman, H. L. Oppenheimer & K. M. Bustamente 22928 Maui US 6/21/2012
21 Coleus scutellarioides (L.) Benth. Along Stainback Highway near junction with Highway 11. PRESERVED_SPECIMEN Herbst, D. R. 926 Hawaii US 2/16/1968
22 Coleus scutellarioides (L.) Benth. Nahiku PRESERVED_SPECIMEN Degener, O. 12462 Maui US 8/30/1939
23 Coleus scutellarioides (L.) Benth. Halelea District; Limahuli Valley; National Tropical Botanical Garden PRESERVED_SPECIMEN Flynn, T. W. 3765 Kauai US 2/15/1990
24 Coleus scutellarioides (L.) Benth. kapa`a, kanepoonui road off kawaihau road. PRESERVED_SPECIMEN n. tangalin 1318 Kauai US 5/29/2007
25 Coleus scutellarioides (L.) Benth. Along Stainback Hwy near junction with Hwy 11. PRESERVED_SPECIMEN Herbst, D. R. 925 Hawaii US 2/16/1968
26 Coleus scutellarioides (L.) Benth. honolulu PRESERVED_SPECIMEN w. t. brigham s.n. Oahu US 1890-08-09
27 Coleus scutellarioides (L.) Benth. Hamakua District, Hana, along Olopawa road to cinder pit. PRESERVED_SPECIMEN Perlman, S. P.; Oppenheimer, H. L.; Bustamente, K. M. 22928 Maui US 6/21/2012
28 Coleus scutellarioides (L.) Benth. Kapa`a, Kanepoonui Road off Kawaihau Road. PRESERVED_SPECIMEN Tangalin, N. 1318 Kauai US 5/29/2007
29 Coleus scutellarioides (L.) Benth. along stainback hwy near junction with hwy 11. PRESERVED_SPECIMEN d. r. herbst 925 Hawaii US 2/16/1968
30 Coleus scutellarioides (L.) Benth. Stainback Hwy. near jct. with Hwy. 11. PRESERVED_SPECIMEN Herbst, D. R. 925 Hawaii US 2/16/1968
31 Coleus scutellarioides (L.) Benth. nahiku PRESERVED_SPECIMEN o. degener 12462 Maui US 8/30/1939
32 Coleus scutellarioides (L.) Benth. Along Stainback Highway near junction with Highway 11. PRESERVED_SPECIMEN Herbst, D. R. 925 Hawaii US 2/16/1968
33 Coleus scutellarioides (L.) Benth. along stainback highway near junction with highway 11. PRESERVED_SPECIMEN d. r. herbst 926 Hawaii US 2/16/1968
34 Coleus scutellarioides (L.) Benth. Hanalei, Wainiha Power House Road. Next to Power House PRESERVED_SPECIMEN Beck, H. T.; Jackson-Beck, L. A. 1200 Kauai US 8/8/1990
35 Coleus scutellarioides (L.) Benth. halelea district; limahuli valley; national tropical botanical garden PRESERVED_SPECIMEN t. w. flynn 3765 Kauai US 2/15/1990
36 Coleus scutellarioides (L.) Benth. honolulu: manoa valley; university of hawaii manoa campus, courtyard of st.john hall PRESERVED_SPECIMEN j. lau 1175 Oahu US 7/1/1985
37 Coleus scutellarioides (L.) Benth. Stainback Hwy. near jct. with Hwy. 11. PRESERVED_SPECIMEN Herbst, D. R. 926 Hawaii US 2/16/1968
38 Coleus scutellarioides (L.) Benth. On side of Hana Hwy 2 mi from Hana. PRESERVED_SPECIMEN Ishikawa, S. S. 163 Maui US 10/14/1972
39 Coleus scutellarioides (L.) Benth. on side of hana hwy 2 mi from hana. PRESERVED_SPECIMEN s. s. ishikawa 163 Maui US 10/14/1972
40 Coleus scutellarioides (L.) Benth. Hanalei Valley, Ohiki Rd. PRESERVED_SPECIMEN Flynn, T. W. 1608 Kauai US 3/24/1986
41 Coleus scutellarioides (L.) Benth. along stainback highway near junction with highway 11. PRESERVED_SPECIMEN d. r. herbst 925 Oahu US 2/16/1968
42 Coleus scutellarioides (L.) Benth. stainback hwy. near jct. with hwy. 11. PRESERVED_SPECIMEN d. r. herbst 925 Hawaii US 2/16/1968
43 Coleus scutellarioides (L.) Benth. sunset beach PRESERVED_SPECIMEN afualo, d. collector number: 125 Oahu US 3/25/1975
44 Coleus scutellarioides (L.) Benth. PRESERVED_SPECIMEN o. degener 12462 Oahu US 8/30/1939
45 Coleus scutellarioides (L.) Benth. East Maui, Hana District, Kalahu, PTBG Kahanu Garden, around Piilanihale Heiau remnant Pandanus forest PRESERVED_SPECIMEN Flynn, T. 2473 Maui BISH 10/20/1987
46 Coleus scutellarioides (L.) Benth. Stainback Hwy. near jct. with Hwy. 11 PRESERVED_SPECIMEN Herbst, D.R. 925 Hawaii BISH 2/16/1968
47 Coleus scutellarioides (L.) Benth. Hiaku Stairs, base of stairs near road PRESERVED_SPECIMEN Wagner, W. L.; Miller, S. W.; Wier, D. B. 5643 Oahu US 6/2/1985
48 Coleus scutellarioides (L.) Benth. Stainback Hwy. near jct. with Hwy. 11 PRESERVED_SPECIMEN Herbst, D.R. 926 Hawaii BISH 2/16/1968
49 Coleus scutellarioides (L.) Benth. 264 california ave., wahiawa red clay PRESERVED_SPECIMEN elia, h.v.k. collector number: 23 Oahu BISH 4/4/1971
50 Coleus scutellarioides (L.) Benth. Honolulu, Nuuanu, Kuakaha open garden bed PRESERVED_SPECIMEN Neal, M.C. s.n. Oahu BISH 6/12/1941
51 Coleus scutellarioides (L.) Benth. nahiku Naturalized over ca. 1 acre in koa tree fern forest on tree trunks and fallen logs. PRESERVED_SPECIMEN degener, o. collector number: 12462 Maui BISH 8/30/1939
52 Coleus scutellarioides (L.) Benth. UH campus, courtyard of St.John Hall bed in full sun PRESERVED_SPECIMEN Lau, J. 1175 Oahu BISH 7/1/1985
53 Coleus scutellarioides (L.) Benth. Nahiku rainy, shrubby aa lava PRESERVED_SPECIMEN Degener, O. 12462 Maui BISH 8/30/1939
54 Coleus scutellarioides (L.) Benth. stainback hwy. near jct. with hwy. 11 PRESERVED_SPECIMEN herbst, d.r. collector number: 925 Hawaii BISH 2/16/1968
55 Coleus scutellarioides (L.) Benth. stainback hwy. near jct. with hwy. 11 PRESERVED_SPECIMEN herbst, d.r. collector number: 926 Hawaii BISH 2/16/1968
56 Coleus scutellarioides (L.) Benth. honolulu: manoa valley; university of hawaii manoa campus, courtyard of st.john hall bed in full sun PRESERVED_SPECIMEN lau, j. collector number: 1175 Oahu BISH 7/1/1985
57 Coleus scutellarioides (L.) Benth. oahu PRESERVED_SPECIMEN brigham, w.t. Oahu BISH 1890-08-08
58 Coleus scutellarioides (L.) Benth. hana, along olopawa road to cinder pit PRESERVED_SPECIMEN perlman, s.p. oppenheimer, h. and bustamente, k. collector number: 22928 Maui BISH 6/21/2012
59 Coleus scutellarioides (L.) Benth. hiaku stairs, base of stairs near road PRESERVED_SPECIMEN w. l. wagner, s. w. miller & d. wier 5643 Oahu US 6/2/1985
60 Coleus scutellarioides (L.) Benth. ahupuaa of halepuaa in the puna district. the plant was collected on the side of the puna trail The plant was growing in an area densely shaded by large Mango trees. The soil is mostly loose cinder. PRESERVED_SPECIMEN yoshida, l. collector number: 79.103 Hawaii BISH 4/16/1979
61 Coleus scutellarioides (L.) Benth. Hilo garden PRESERVED_SPECIMEN Fosberg, F.R. 10266 Hawaii BISH 9/6/1933
62 Coleus scutellarioides (L.) Benth. Honolulu: Manoa Valley; University of Hawaii Manoa Campus, courtyard of St.John Hall bed in full sun PRESERVED_SPECIMEN Lau, J. 1175 Oahu BISH 7/1/1985
63 Coleus scutellarioides (L.) Benth. Laie, Temple loam PRESERVED_SPECIMEN Ziegelbauer, K. 32 Oahu BISH 3/13/1975
64 Coleus scutellarioides (L.) Benth. Halelea, Limahuli; NTBG PRESERVED_SPECIMEN Flynn, T. 3765 Kauai BISH 2/15/1990
65 Coleus scutellarioides (L.) Benth. Hilo PRESERVED_SPECIMEN Faurie, A. 911 Hawaii BISH 5/1/1909
66 Coleus scutellarioides (L.) Benth. ahupuaa of Halepuaa in the Puna District, on the side of the Puna Trail The plant was growing in an area densely shaded by large Mango trees. The soil is mostly loose cinder. PRESERVED_SPECIMEN Yoshida, L. 79.103 Hawaii BISH 4/16/1979
67 Coleus scutellarioides (L.) Benth. hanalei valley, ohiki rd pasture PRESERVED_SPECIMEN flynn, t. collector number: 1608 Kauai BISH 3/24/1986
68 Coleus scutellarioides (L.) Benth. East Maui, side of Hana Hwy 2 mi from Hana ferns and grasses PRESERVED_SPECIMEN Ishikawa, S. 163 Maui BISH 10/14/1972
69 Coleus scutellarioides (L.) Benth. halelea, limahuli; ntbg Wet 'ohi'a forest with other native trees and a treefern and native shrub understory. PRESERVED_SPECIMEN flynn, t. collector number: 3765 Hawaii BISH 2/15/1990
70 Coleus scutellarioides (L.) Benth. pupukea; boys campus Rocky south rim. PRESERVED_SPECIMEN utai, m. collector number: 8 Oahu BISH 2/1/1975
71 Coleus scutellarioides (L.) Benth. 264 California Ave., Wahiawa red clay PRESERVED_SPECIMEN Elia, H.V.K. 23 Oahu BISH 4/4/1971
72 Coleus scutellarioides (L.) Benth. honolulu, tantalus, puu ualakaa gate wet forest, roadside; with Eucalyptus citriodora, Toona ciliata, Setaria gracilis PRESERVED_SPECIMEN ray, g. gibney, e. collector number: 42 Oahu BISH 8/27/1997
73 Coleus scutellarioides (L.) Benth. Honolulu, Tantalus, Puu Ualakaa gate wet forest, roadside; with Eucalyptus citriodora, Toona ciliata, Setaria gracilis PRESERVED_SPECIMEN Ray, G. Gibney, E. Collector Number: 42 Oahu BISH 8/27/1997
74 Coleus scutellarioides (L.) Benth. Pupukea; Boys Campus clay silt PRESERVED_SPECIMEN Utai, M. 8 Oahu BISH 2/1/1975
75 Coleus scutellarioides (L.) Benth. East Maui, 15 mi W of Hana along Hwy. 36 with Eucalyptus, Monstera PRESERVED_SPECIMEN Henrickson, J.S. 3723 Maui BISH 6/26/1969
76 Coleus scutellarioides (L.) Benth. ahupuaa of Halepuaa in the Puna District. THe plant was collected on the side of the Puna Trail The plant was growing in an area densely shaded by large Mango trees. The soil is mostly loose cinder. PRESERVED_SPECIMEN Yoshida, L. Collector Number: 79.103 Hawaii BISH 4/16/1979
77 Coleus scutellarioides (L.) Benth. stainback hwy. near jct. with hwy. 11 on muddy rock PRESERVED_SPECIMEN herbst, d.r. collector number: 925 Kauai BISH 2/16/1968
78 Coleus scutellarioides (L.) Benth. ahupuaa of Halepuaa in the Puna District, on the side of the Puna Trail The plant was growing in an area densely shaded by large Mango trees. The soil is mostly loose cinder. PRESERVED_SPECIMEN Yoshida, L. Collector Number: 79.103 Hawaii BISH 4/16/1979
79 Coleus scutellarioides (L.) Benth. east maui, hana district, kalahu, ptbg kahanu garden, around piilanihale heiau remnant Pandanus forest PRESERVED_SPECIMEN flynn, t. collector number: 2473 Maui BISH 10/20/1987
80 Coleus scutellarioides (L.) Benth. ahupuaa of halepuaa in the puna district, on the side of the puna trail The plant was growing in an area densely shaded by large Mango trees. The soil is mostly loose cinder. PRESERVED_SPECIMEN yoshida, l. collector number: 79.103 Hawaii BISH 4/16/1979
81 Coleus scutellarioides (L.) Benth. ahupuaa of Halepuaa in the Puna District. THe plant was collected on the side of the Puna Trail The plant was growing in an area densely shaded by large Mango trees. The soil is mostly loose cinder. PRESERVED_SPECIMEN Yoshida, L. 79.103 Hawaii BISH 4/16/1979
82 Coleus scutellarioides (L.) Benth. East Maui, Hana Rd. near Hana PRESERVED_SPECIMEN Fosberg, F.R. 9910 Maui BISH 8/20/1933
83 Coleus scutellarioides (L.) Benth. laie PRESERVED_SPECIMEN guinto, j. collector number: 22 Oahu BISH 3/6/1986
84 Coleus scutellarioides (L.) Benth. hanalei, wainiha power house road. next to power house Planted along stream. PRESERVED_SPECIMEN h. t. beck & l. jackson-beck 1200 Oahu BISH 8/8/1990
85 Coleus scutellarioides (L.) Benth. hilo garden PRESERVED_SPECIMEN fosberg, f.r. collector number: 10266 Hawaii BISH 9/6/1933
86 Coleus scutellarioides (L.) Benth. Hanalei Valley, Ohiki Rd pasture PRESERVED_SPECIMEN Flynn, T. 1608 Kauai BISH 3/24/1986
87 Coleus scutellarioides (L.) Benth. Hilo PRESERVED_SPECIMEN Faurie, A. Collector Number: 911 Hawaii BISH 5/1/1909
88 Coleus scutellarioides (L.) Benth. hilo PRESERVED_SPECIMEN faurie, a. collector number: 911 Maui BISH 5/1/1909
89 Coleus scutellarioides (L.) Benth. ahupuaa of Halepuaa in the Puna District. THe plant was collected on the side of the Puna Trail Mesic wet forest, disturbed roadside PRESERVED_SPECIMEN G. J. Ray 42 Hawaii HAPI 8/27/1997
90 Coleus scutellarioides (L.) Benth. uh campus, courtyard of st.john hall bed in full sun PRESERVED_SPECIMEN lau, j. collector number: 1175 Oahu BISH 7/1/1985
91 Coleus scutellarioides (L.) Benth. Honolulu, Tantalus, Puu Ualakaa gate PRESERVED_SPECIMEN S. P. Perlman, H. L. Oppenheimer & K. M. Bustamente 22928 Oahu HAPI 6/21/2012
92 Coleus scutellarioides (L.) Benth. east maui, hana rd. near hana PRESERVED_SPECIMEN fosberg, f.r. collector number: 9910 Maui BISH 8/20/1933
93 Coleus scutellarioides (L.) Benth. east maui, 15 mi. w of hana along hwy. 36 Dense tree 15' ht, off trail on side of valley; under large Acacia koa and Antidesma platyphyllum PRESERVED_SPECIMEN henrickson, j.s. collector number: 3723 Oahu BISH 6/26/1969
94 Coleus scutellarioides (L.) Benth. laie, temple loam PRESERVED_SPECIMEN ziegelbauer, k. collector number: 32 Oahu BISH 3/13/1975
95 Coleus scutellarioides (L.) Benth. Hilo garden PRESERVED_SPECIMEN Fosberg, F.R. Collector Number: 10266 Hawaii BISH 9/6/1933
96 Coleus scutellarioides (L.) Benth. east maui, side of hana hwy 2 mi. from hana ferns and grasses PRESERVED_SPECIMEN ishikawa, s. collector number: 163 Maui BISH 10/14/1972
97 Coleus scutellarioides (L.) Benth. East Maui, Hana, along road to dump. Growing in weedy brush along road. PRESERVED_SPECIMEN Hobdy, R.W. 3221 Maui BISH 7/14/1990
98 Coleus scutellarioides (L.) Benth. Hana, along Olopawa road to cinder pit PRESERVED_SPECIMEN Perlman, S. 22928 Maui BISH 6/21/2012
99 Coleus scutellarioides (L.) Benth. honolulu, nuuanu, kuakaha open garden bed PRESERVED_SPECIMEN neal, m.c. Oahu BISH 6/12/1941
100 Coleus scutellarioides (L.) Benth. Sunset Beach silty loam PRESERVED_SPECIMEN Afualo, D. 125 Oahu BISH 3/25/1975
101 Coleus scutellarioides (L.) Benth. Honolulu, Tantalus, Puu Ualakaa gate wet forest, roadside; with Eucalyptus citriodora, Toona ciliata, Setaria gracilis PRESERVED_SPECIMEN Ray, G. 42 Oahu BISH 8/27/1997